Photo of symptoms of crumpled leaf on bean leaf
Foliar chlorosis and necrosis resulting from a genetic disorder called crumpled leaf, a genetic trait called crumpled leaf or LCR caused by an incompatible developmental reaction when beans from different centers of domestication are crossed, e.g., crosses of Mesoamerican bush blue lake materials with Andean Midwestern types of beans. Symptoms can resemble those caused by virus infections. Lines with the trait may vary in intensity of expression, and the expression can vary over the season.

Photo Source: Lindsey du Toit, Washington State University